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In all drawings, the distal pole is on top, the proximal pole at the bottom, unless otherwise specified.
e = equatorial view, p = polar view; H = high level, L = low level.
The terms printed in bold are preferred.
The colors are according the scheme described in the
introduction.
Click on a drawing for a full sized picture.
Decussate tetrad (Walker and Doyle, 1975) | ||
A
multiplanar tetrad of
pollen grains or spores
arranged in two pairs lying across one another, the pairs (dyads) more or less at right angles to each other. Example: Orophea
(Annonaceae). |
Demicolpus (pl. demicolpi, adj. demicolpate) (Erdtman, 1952) | ||
An ectocolpus divided into two parts. Example: Amylotheca
(Loranthaceae).
|
Di- | ||
A prefix for two.
|
Dicolpate, dicolporate, diporate (adj.) (Iversen and Troels-Smith, 1950) | ||
Describing
pollen grains with two
ectocolpi, two
compound apertures or two
pores.
|
Dicolporate (adj.) (Cranwell, 1953) | ||
Synonym of
diploporate.
|
Digitate (adj.) (Skvarla and Larson, 1965) | ||
With
bacula or
columellae which are branched distally into two or more parts. Example:
Polygonum bistorta. |
Diorate (adj.) (Erdtman, 1952) | ||
Synonym of
diploporate. |
Diploporate (adj.) (Fægri and Iversen, 1964) | ||
Describing an
ectocolpus with two
endoapertures. Example: Didymeles (Didymelaceae).
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Diploxylonoid (adj.) (Traverse, 1988) | ||
Describing
bisaccate pollen
grains in which the outline of the sacci in polar view is discontinuous with the outline of the corpus so that the grains seem to consist of three distinct, more
or less oval parts.
|
Discordant pattern (Fægri and Iversen, 1989) | ||
A pattern in a tectate
pollen grain in which the arrangement of the
columellae is different from that of the elements on the tectum. Example: Geranium (Geraniaceae).
|
Dispersal unit | ||
The morphological unit in which mature pollen grains or spores are shed, which may range from individuals (monads), to pairs (dyads), groups of four (tetrads), or groups of more than four (polyads). Larger, indeterminate numbers of pollen grains or spores may also be dispersed as pollinia or massulae. |
Dissections (Couper and Grebe, 1961) | ||
Rounded to elongated cavities in a cingulum or
zona. Example: Vallatisporites ciliaris.
|
Distal (adj.) | ||
A common descriptive term (Jackson, 1928) used in contrast to proximal, applied in
palynology to features on the surface that face
outward in the tetrad stage (Wodehouse, 1935).
|
Distal face (Erdtman, 1952) | ||
That part of a
palynomorph that faces outwards the centre of the tetrad, between
equator and distal
pole.
|
Distal pole (Erdtman, 1952) | ||
The centre of the surface of the
distal face.
|
Disulcate (adj.) (Harley, 1998) | ||
Describing pollen grains with sulci arranged in pairs. Two
types of disulcate pollen are distinguished: equatorial disulcate, with opposing, eaquatorially
arranged sulci and distal disulcate with paired sulci lying parallel to the long axis of the pollen
grain on the distal face. Examples: Metroxylon salomonense (Palmae) (equatorial disulcate),
Chamaerops humilis (Palmae) (distal disulcate).
|
Duplibaculate (adj.) (Erdtman, 1952) | ||
Synonym of
duplicolumellate. |
Duplicolumellate (adj.) (Reitsma, 1970) | ||
With
columellae in two rows under each
murus. Example: Polygonum persicaria (Polygonaceae).
|
Dyad | ||
General term for two
microspores (pollen
grains or spores) united as a dispersal
unit.
|
Echinate (adj.) (sing. echina, pl. echinae) (Wodehouse, 1928) | ||
Describing
pollen and spores with
an ornamentation comprising
spines longer than 1 µm.
|
Echinolophate (adj.) (Wodehouse, 1928) | ||
Describing a
lophate pollen grain
with echinate ridges.
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Ectexine (sensu Fægri, 1956) | ||
The outer part of the
exine, which stains positively with basic fuchsin in optical microscopy and has
higher electron density in conventionally prepared TEM sections. Orthographical variant:
ektexine.
|
Ecto- | ||
A prefix for outer. |
Ectoaperture (Van Campo, 1958) | ||
An
aperture in the outer layer of the
sporoderm.
|
Ectointine (Freytag, 1968) | ||
Synonym of exintine. |
Ektannulus (pl. ektannuli) (Batten and Christopher, 1981) | ||
An abrupt thickening of the outer wall layer in the region
of the exogerminal. Example: Krutzschipollis.
|
Ektexine (Erdtman, 1943) | ||
Orthographic variant of
ectexine.
|
Ekto- | ||
Orthographical variant of the prefix ecto-. |
Elater | ||
A spirally twisted, filamentous band attached to certain
spores and some fossil
pollen grains (Jackson, 1928).
Example: Equisetum (Equisetaceae). |
Endannulus (pl. endannuli) (Thomson and Pflug, 1953) | ||
An
annulus formed by the
endexine of a pollen
grain. Example: Atlantopollis.
|
Endexine (sensu Fægri, 1956) | ||
The inner part of the
exine which remains relatively unstained with basic fuchsin in optical microscopy
and has a lower electron density in conventionally prepared TEM sections.
|
Endintine (Kress and Stone, 1982) | ||
The inner, cellulosic zone of the
intine which is adjacent to the cytoplasm and in fluorescence microscopy stains
positively with PAS calcofluor.
|
Endo- | ||
A prefix for inner. |
Endoaperture (Van Campo, 1958) | ||
An
aperture in the inner layer of the
sporoderm, often the inner aperture of a
compound aperture.
|
Endoaperture area (Verbeek-Reuvers, 1976) | ||
The region of the
nexine of a single
compound aperture which encloses a number of endoapertures.
|
Endocingulum (pl. endocingula, adj. endocingulate) (Reitsma, 1966) | ||
A ring-shaped
endoaperture continuous around a
pollen grain and lying in the
equatorial plane. Example: Polygonum convolvulus
(Polygonaceae).
|
Endocrack (Oldfield, 1959) | ||
An irregular groove occurring in the inner surface of the
nexine/endexine and readily apparent in acetolysed
pollen. Example: Anemone nemorosa
(Ranunculaceae). |
Endogerminal (adj.) (Batten and Christopher, 1981) | ||
Describing an
aperture in the inner wall layer.
|
Endointine (Freytag, 1968) | ||
Orthographic variant of endintine. |
Endoplica (pl. endoplicae) (Thomson and Pflug, 1953) | ||
A fold or swelling of the inner wall layer.
|
Endosculpture (Van Campo, 1971) | ||
Sculpturing occurring on the inner surface of the nexine/endexine. |
Endospore | ||
The innermost layer of a
spore wall (Jackson, 1928).
|
Endosporium (Erdtman, 1943) | ||
Synonym of
intine. |
Equator (Wodehouse, 1935) | ||
The dividing line between the
distal and proximal
faces of a pollen grain or spore.
|
Equatorial axis | ||
Often misappropriately used as a synonym of equatorial diameter. |
Equatorial bridge (Moore and Webb, 1978) | ||
Synonym of
bridge. |
Equatorial diameter (Erdtman, 1943) | ||
A line, lying in the
equatorial plane, perpendicular to the
polar axis and passing through it.
|
Equatorial lacuna (pl. equatorial lacunae) (Wodehouse, 1928) | ||
A lacuna situated on the equator in the mesocolpial region of a
lophate pollen grain.
Example: Tragopogon pratensis (Compositae).
|
Equatorial limb | ||
Synonym of
equatorial outline. |
Equatorial outline | ||
General description of the equator when a pollen grain is seen in
polar view.
|
Equatorial plane (Fægri and Iversen, 1950) | ||
The plane perpendicular to the
polar axis and lying midway between the
poles. |
Equatorial ridge (Wodehouse, 1928) | ||
A ridge lying along the
equator in the mesocolpial region, usually applied to
lophate pollen grains.
Example: Cichorium intybus (Compositae). |
Equatorial view (Erdtman, 1943) | ||
The view of a
pollen grain or spore
where the equatorial plane is directed towards
the observer.
|
Equiaxe (adj.equiaxal) (Van Campo, 1966) | ||
Pollen grains with a polar
axis equal to the equatorial
diameter.
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Erect (adj.) (Reitsma, 1970) | ||
Synonym of
prolate. |
Etectate (adj.) (Walker, 1976) | ||
Describing
pollen grains interpreted as having lost their tectum during
evolutionary development.
|
Eu- | ||
A prefix for true.
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Euintine (Kuprianova, 1956) | ||
Synonym of endintine. |
Eurypalynous (adj.) (Erdtman, 1952) | ||
Describing plant taxa characterized by possession of a great
diversity of palynomorphs.
|
Eutectum (adj.eutectate) | ||
Describing a
pollen grain with a continuous
tectum.
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Exine (pl. exines, adj. exinal, exinous) (Fritzsche, 1837) | ||
The outer layer of the wall of a
palynomorph, which is highly resistant to strong acids and bases, and is
composed primarily of sporopollenin. |
Exine 1 (Erdtman, 1969) | ||
Synonym of ectexine. |
Exine 2 (Erdtman, 1969) | ||
Synonym of endexine. |
Exintine (Kress and Stone, 1982) | ||
An outer (pectic) layer of the
intine, lying below the
nexine, and which stains positively with alcian blue.
|
Exitus (Wodehouse, 1935) | ||
The site of exit of the
pollen tube from the
sporoderm.
|
Exoexine (Potonié, 1934) | ||
Synonym of
sexine. |
Exogerminal (adj.) (Batten and Christopher, 1981) | ||
Describing an
aperture formed in the outer wall layer of the
exine.
|
Exospore | ||
The outer layer of a
spore wall (Jackson, 1928).
|
Exosporium (Erdtman, 1943) | ||
Synonym of exine. |
Fastigium (pl. fastigia, adj. fastigiate) (Reitsma, 1966) | ||
Cavity in a
colporate grain, appearing as a separation of the inner part of the exine from the domed sexine in the region of the endoaperture. Example: Hypericum
(hypericaceae).
|
Fenestrate (adj.) (Iversen and Troels-Smith, 1950) | ||
Describing a class of
pollen grains characterized by large, window-like spaces lacking a tectum.
|
Fimbria (pl. fimbriae, adj. fimbriate) | ||
Long, hair-like appendages (Jackson, 1928).
|
Fischer's law/rule (Erdtman, 1952) | ||
The name given to the widespread arrangement in
developmental tetrads of tri-aperturate pollen grains whereby the
apertures form in pairs at six points in the tetrad. Example: Ericaceae
|
Fissura (pl. fissurae) (Potonié, 1934) | ||
A sharp, straight split that appears during germination in certain inaperturate pollen grains. Examples: Taxodium (Taxodiaceae), Cupressus (Cupressaceae). |
Flange | ||
A general term, used to describe equatorial extensions of
spores (Jackson, 1928).
|
Foot layer (Fægri, 1956) | ||
The inner layer of the ectexine.
|
Foramen (pl. foramina, adj. forate, which is an abbreviation of foraminate) (Erdtman, 1952) | ||
Synonym of
pore.
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Fossaperturate (adj.) (Erdtman, 1952) | ||
Refers to an equatorially aperturate, lobate pollen grain with the
apertures in the indentations between the lobes.
|
Fossula (pl. fossulae, adj. fossulate) (Fægri and Iversen, 1950) | ||
A feature of
ornamentation consisting of an elongated, irregular groove in the surface.
|
Fossula (Kuprianova, 1948) | ||
Synonym of
areola. |
Foveola (pl. foveolae, adj. foveolate) (Erdtman, 1952) | ||
A feature of
ornamentation consisting of more or less rounded depressions or
lumina more than 1µm in diameter. The distance between foveolae is greater than their
breadth.
|
Frustillum (pl. frustilla, adj. frustillate) (Fægri and Iversen, 1964) | ||
Synonym of
areola. |
Furrow | ||
A common word for an elongate
aperture.
|
Furrow membrane (Wodehouse, 1935) | ||
Synonym of
colpus membrane. |
Galea (pl. galeae, adj. galeate) (Sullivan, 1964) | ||
A relatively large element of the outer wall of a
spore consisting of a sharply tapering spine and a broad
bulbous base. |
Garside's law/rule (Garside, 1946; Erdtman, 1952) | ||
The name given to an unusual arrangement in
developmental tetrads of tri-aperturate pollen whereby
the apertures form in groups of three at four points in the
tetrad.
|
Geminicolpate (adj.) (Erdtman, 1952) | ||
Describing
pollen grains with colpi
arranged in pairs.
|
Gemma (pl. gemmae, adj. gemmate) (Iversen and Troels-Smith, 1950) | ||
A sexine element which is constricted at its base, higher than 1µm, and that has approximately the same width as its height. |
Generative cell | ||
The cell in a pollen grain which divides to form male gametes (Jackson, 1928). |
Geniculum (pl. genicula, adj. geniculate) (Potonié, 1934) | ||
A bulge in the equatorial exine of the colpus, often associated with a separation of the sexine from the
nexine and the rupturing of the latter. Examples: Fraxinus (Olea
ceae), Quercus (Faga- ceae).
|
Germinal aperture (Wodehouse, 1935) | ||
A hole in the furrow membrane through which the
pollen tube emerges.
|
Goniotreme (adj.) (Erdtman and Straka, 1961) | ||
Synonym of
angulaperturate. |
Granular exine (Van Campo and Lugardon, 1973) | ||
A type of exine stratification in which the infratectal layer is composed of more or less rounded,
granules rather than of
columellae or other structures.
|
Granulate granulatus (adj.) (Erdtman, 1947) | ||
With granules. |
Granule (pl. granules, adj. granular, granulose) | ||
General word for a small, rounded element. |
Granulum (pl.granula, adj.granulate, granulose) (sensu Erdtman, 1952) | ||
A very small and rounded element of the sexine/ectexine that is less than 1µm in all directions.
|
Granum (pl. grana) (Potonié, 1934) | ||
The combined elements gemmae and verrucae. |
Groove | ||
A general descriptive word.
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Gula (pl. gulae, adj. gulate) (Potonié and Kremp, 1955) | ||
A rather ornate projecting, neck-like, extension on the
proximal face of a
trilete spore. Example:
Lagenicula horrida.
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Halo (Erdtman, 1952; Fægri and Iversen, 1989) | ||
A clear zone around a well defined feature such as a
spine or an
aperture. Examples: Ranunculus acris (Ranunculaceae),
Valeriana (Valerianaceae). |
Hamulate (adj.) (Krutzsch, 1959) | ||
Describing a form of
rugulate
ornamentation consisting of irregularly arranged, winding, or angular
rounded muri of varying thickness, which do not form
a distinct reticulum, but rather a maze-like pattern.
Example: Lycopodiella inundata (Lycopodiaceae).
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Haploxylon-type (Rudolph, 1935) | ||
Bisaccate pollen in
which the outline of the sacci in polar view is more or less continuous with the outline of the
corpus, so that the grains appear a more or less
smooth ellipsoidal form. Examples: Pinus cembra, Picea (Pinaceae).
|
Haploxylonoid (adj.) (Traverse, 1988) | ||
Describing
bisaccate pollen in
which the outline of the sacci in polar view is more
or less continuous with the outline of the corpus, so
that the grains appear a more or less smooth ellipsoidal form.
|
Harmomegathy (adj. harmomegathic) (Wodehouse, 1935) | ||
The process by which pollen grains and spores change in shape to accommodate variations in the volume of the cytoplasm caused by changing hydration. |
H-endoaperture (Punt and Nienhuis, 1976) | ||
An elaborate endoaperture, consisting of a central part which connects two lateral, longitudinal elongations, forming an "H" shape. Examples: Cornus (Cornacaea), Centaurium (Gentianaceae). |
Heterobrochate (adj.) (Erdtman, 1952) | ||
Describing a
reticulum with brochi of different sizes. Examples:
Adoxa Moschatelina (Adoxaceae), Forsythia europaea (Oleaceae).
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Heterocolpate (adj.) (Iversen and Troels-Smith, 1950) | ||
Describing
pollen grains with both simple and
compound colpi present. Examples: Lythrum,
Peplis (Lythraceae), Myosotis Boraginaceae).
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Heteropolar (adj.) (Erdtman, 1952) | ||
Describing
pollen or spores in
which the distal and
proximal faces of the
exine are different, either in shape,
ornamentation or apertural system. Example: Echium
vulgare (Boraginaceae).
|
Heterosporous (adj.) | ||
Describing plants producing both
microspores and
megaspores (Jackson, 1928).
|
Hilum (pl. hila, adj. hilate) (Erdtman, 1952) | ||
Circular, indistinctly delimited, irregular aperture or thinning in
spores. Examples: Aequitriradites verrucosus, Couperisporites
tabulatus.
|
Homobrochate (adj.) (Erdtman, 1952) | ||
Describing a
reticulum with brochi of the same sizes. Example:
Armeria maritima (Plumbaginaceae).
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Homosporous (adj.) | ||
Describing plants producing only one kind of spores (Jackson, 1928).
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Horn (Huynh, 1970) | ||
An elongated part of an
endoaperture, which is curved towards one of the poles. Example: Anagallis arvensis (Primulaceae).
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