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In all drawings, the distal pole is on top, the proximal pole at the bottom, unless otherwise specified.
e = equatorial view, p = polar view; H = high level, L = low level.
The terms printed in bold are preferred.
The colors are according the scheme described in the
introduction.
Click on a drawing for a full sized picture.
S-type tetrad (Moar, 1993) | ||
A tetrad in which only one members is fully developed. Example:
Leucopogon fasciculatus (Epacridaceae).
|
Saccoid (adj.)(Brugman, 1983) | ||
A saccus-like expansion of the
exine with a complex infrastructure build up by a three
dimensional network of sexine elements, extending
to and fused with the nexine.
|
Saccus (pl. sacci, adj. saccate) (Erdtman, 1952) | ||
A sac formed by an expansion of the exine of a
pollen grain and at least partly filled with an
alveolate
infrastructure.
|
Scabrate (adj.) (sing. scabra, pl. scabrae) (Iversen and Troels-Smith, 1950) | ||
Describing elements of ornamentation, of any shape,
smaller than 1µm in all directions. Examples: Quercus (Fagaceae), Artemisia
(Compositae).
|
Sclerine (Erdtman, 1952) | ||
A term encompassing both
exine and perine that
can be used whether a perine is present or not; sporoderm excluding the
intine.
|
Scrobiculus (pl. scrobiculi, adj. scrobiculate) (Potonié, 1934) | ||
Synonym of
punctum. |
Sculptine (Erdtman, 1948) | ||
A term encompassing both
exine and perine
(sclerine) but excludes the nexine
and so provides a neutral term for the sculptured layer when there is doubt whether the pattern belongs to the
exine or the perine.
|
Sculpural density (Balme, 1988) | ||
The estimated number of sculptural elements in an area of 100 µm2 of the surface of the exine. |
Sculpture (Kuprianova, 1948) | ||
Orthogriphical variant of sculpturing. |
Sculpturing (adj. sculptured) (Potonié, 1934) | ||
The surface relief, or topography, of a pollen grain or spore.
|
Seed- megaspore | ||
A large, functional megaspore associated with three small, presumably aborted spores present in some fossil lycopsids. Example: Cystosporites. |
Semi-erect (adj.) (Reitsma, 1970) | ||
Synonym of subprolate. |
Semitectum (adj. semitectate) (Fægri and Iversen, 1964) | ||
A partially discontinuous tectum in which
the tectal perforations are equal to or wider than the
muri and usually larger than 1µm in diameter.
|
Semitransverse (adj.) (Reitsma, 1970) | ||
Synonym of suboblate. |
Sexine (Erdtman, 1952) | ||
The outer, sculptured layer of the exine, which lies above the nexine.
|
Sexine 1, 2, 3, 4, 5 (Reitsma, 1970) | ||
A system of sexine stratification in which sexine 1 is the innermost and sexine 5 in the
outermost layer of the sexine.
|
Shape classes (pl.) (Erdtman, 1943) | ||
Categories of
pollen and spore shape
based on the relations between polar axis (P) and
equatorial diameter (E).
|
Simplibaculate (adj.) (Erdtman, 1952) | ||
Synonym of
simplicolumellate. |
Simplicolumellate (adj.) (Reitsma, 1970) | ||
With a single row of
columellae under each
murus. Example: Viburnum opulus (Caprifoliaceae).
|
Sinu-aperturate (adj.) (Erdtman, 1952) | ||
Describing a
pollen grain in which the equatorial apertures are situated in the middle of
concave sides when seen in polar view.
|
Sole (APLF, 1975) | ||
Synonym of
foot layer. |
Special wall (Beer, 1911) | ||
Pollen mother-cell wall |
Spheroidal (Erdtman, 1943) | ||
Describing the shape of a
pollen grain or spore
in which the polar axis and the equatorial diameter are approximately equal.
|
Spina (pl.spinae) (Potonié, 1934) | ||
Synonym of Spine. |
Spine (adj. spiny/spinose) (Erdtman, 1952) | ||
A general word, applied in
palynology to long and tapering pointed elements, exceeding 1µm.
|
Spinule (adj. spinulose) (Erdtman, 1952) | ||
Small spines, less than 3µm in length.
|
Spiraperturate (adj.) (Erdtman, 1952) | ||
Describing a
pollen grain with one or more spiral apertures. Example: Eriocaulon
aquaticum (Eriocaulaceae).
|
Spore | ||
A general term for the usually microscopic, unicellular,
asexual or sexual reproductive units of cryptogams and fungi (Jackson,
1928).
|
Sporoderm (Bischoff, 1833) | ||
The entire wall of a pollen grain or spore. |
Sporomorph (Erdtman, 1947) | ||
A general term for spore-like palynomorphs. |
Sporopollenin (Zetsche and Vicari, 1931) | ||
The name given to the acetolysis resistant biopolymers which make up most of the material of the exine. |
Square tetrad (Erdtman, 1945b) | ||
Synonym of
tetragonal tetrad. |
Stenopalynous (adj.) (Erdtman, 1952) | ||
Describing plant taxa characterized by only a slight
variation in their palynomorphs.
|
Stephano- | ||
Synonym of zono-.
|
Stratum (APLF, 1975) | ||
A subdivision of a major layer of the sporoderm. |
Striae (pl.) (sing. stria) (Fægri and Iversen, 1950) | ||
Grooves between elongated sculpturing elements. |
Striate (adj.) (Iversen and Troels-Smith, 1950) | ||
A general descriptive term applied in palynology to elongated, generally parallel elements separated
by grooves.
|
Striato-reticulate (adj.) (Erdtman, 1952) | ||
Describing a pattern in which parallel or subparallel
muri are cross-linked to form a reticulum in the grooves. The connections between the muri
lie on a single level or different levels. Example: Gentiana pneumonathe (Gentianaceae). |
Structure (adj. structurate, structured) (Potonié, 1934) | ||
The internal construction of the
pollen or spore wall.
|
Sub- | ||
A prefix for under, or less than. |
Suberect (adj.) (Reitsma, 1970) | ||
Synonym of
prolate spheroidal. |
Subisopolar (adj.) (Walker and Doyle, 1975) | ||
Describing a pollen grain or spore in which the proximal and distal faces are slightly different. Examples: one face is convex and the other is less convex (Banksia, Proteaceae); on one face viscin threads are connected (Oenothera, Onagraceae). |
Sub-layer | ||
Synonym of stratum. |
Suboblate, (adj.) (Erdtman, 1952) | ||
Describing the shape of a
pollen grain or spore
in which the ratio between the polar axis and the
equatorial diameter is 0.75-0.88.
|
Subprolate (adj.) (Erdtman, 1952) | ||
Describing the shape of a
pollen grain or spore
in which the ratio between the polar axis and the
equatorial diameter is 1.14-1.33.
|
Subspheroidal (adj.) (Erdtman, 1952) | ||
Describing the shape of a
pollen grain or spore
in which the ratio between the polar axis and the
equatorial diameter is 0.75-1.33.
|
Subtransverse (adj.) (Reitsma, 1970) | ||
Synonym of
oblate spheroidal. |
Subturma (pl. subturmae) (Potonié, 1956) | ||
A group of form-genera of fossil
spores or pollen in the
turma-system of Potonié.
|
Successiform (Van Campo, 1967) | ||
Referring to a phyletic series of pollen types with increasing numbers of apertures, ranging from tricolpate to pantocolpate and pantoporate. |
Sulculus (pl. sulculi, adj. sulculate) (Erdtman, 1952) | ||
An elongated
latitudinal
ectoaperture not situated at a
pole.
|
Sulcus (pl. sulci, adj. sulcate) (Erdtman, 1952) | ||
An elongated
latitudinal
ectoaperture situated at the
distal or proximal
pole of a pollen grain.
|
Supra-(prefix) | ||
A prefix for above.
|
Supratectal (adj.) (Erdtman, 1969) | ||
Indicating the position of features, such as spines, on the top of the tectum. |
Suture (Potonié and Kremp, 1955) | ||
Synonym of
commissure.
|
Sylvestris-type (Rudolph, 1935) | ||
Bisaccate pollen
grains in which the outline of the sacci in polar view is discontinuous with the outline of the corpus so that the grains seem to consist of three distinct, more
or less oval parts. Examples: Pinus sylvestris, Abies (Pinaceae).
|
Syn- | ||
A prefix indicating the fusion or anastomosis of features. |
Syncolp(or)ate (adj.) (sensu Erdtman, 1952) | ||
Describing a
pollen grain with two or more simple (or
compound)
colpi the ends of which anastomose at the
pole. Example: Primula farinosa (Primulaceae).
|
Synrugoidate (adj.) (Erdtman, 1935; Jalan and Kapil, 1964) | ||
Describing a
pollen grain with six
colpi of which three are long and meeting at one
pole and three are short and not meeting at either pole. Example: Schisandra
grandiflora (Schisandraceae). |
T-shaped tetrad (Walker and Doyle, 1975 ) | ||
A uniplanar tetrad in which two of the members are perpendicular to the
other two so that tetrad has the shape of the letter "T". Example: Typha spp.
(Typhaceae).
|
T-type tetrad (Moar, 1993) | ||
A tetrad in which all four members are fully developed. Example:
Pentachondra pumila (Epacridaceae).
|
Taenia (pl. taeniae, adj. taeniate) (Leschik, 1956) | ||
One or more strap-like, more or less parallel strips of
exine on the
proximal and/or distal
sides of the corpus of certain fossil gymnospermous
pollen grains. Examples: Striatites,
Vittatina.
|
Tectum (pl. tecta, adj. tectate) (Fægri and Iversen, 1950) | ||
The layer of sexine, which forms a roof over the
columellae, granules or other
infratectal elements.
|
Tectum perforatum (adj. tectate perforate) (Iversen and Troels-Smith, 1950) | ||
A tectum with perforations smaller than 1µm in diameter.
|
Tectum imperforatum (adj. tectate imperforate) (Walker and Doyle, 1975) | ||
With a continuous
tectum, without perforations.
|
Tectum solidum (Fægri and Iversen, 1975) | ||
Synonym of
eutectum. |
Tegillum (pl. tegilla, adj. tegillate) (Erdtman, 1952) | ||
Synonym of tectum. |
Tenui- (Erdtman, 1952) | ||
A prefix for thin.
|
Tenuitas (pl. tenuitates) (Potonié, 1934) | ||
A general term for a thinning, that has been applied to
many different situations in palynology.
|
Tetrad | ||
A general term for a group of four united pollen grains or
spores, either as a dispersal
unit or as a developmental stage.
|
Tetrad mark (Couper and Grebe, 1961) | ||
The
monolete or trilete mark on the
proximal face of a
spore or, more rarely, a
pollen grain.
|
Tetrad stage | ||
The period during post-meiotic development when the four
microspores or
megaspores are united by the presence of a temporary special wall.
|
Tetragonal tetrad (Grebe, 1971) | ||
A uniplanar tetrad in which all four members are in contact at the centre
of the tetrad so that, in the correct orientation, the
adjacent walls form a cross. Example: Uvariastrum hexaloboides (Annonaceae)
|
Tetrahedral tetrad (Grebe, 1971) | ||
A
multiplanar tetrad in which each member is in contact with three
others, so that the centers of the grains define a tetrahedron. Example: Erica (Ericaceae).
|
Tilia structure (Praglowski, 1971) | ||
A sexine structure describing
a pertacte tectum
provided with funnel-like concavities of which the bottoms coalesce with a single underlying
columella, to form one unit. Example: Tilia cordata
(Tiliaceae). |
Torus (pl. tori) (Thomson and Pflug, 1953) | ||
An arcuate invagination or protuberance of the exine more or less paralleling the
laesura of a spore in
the interradial area.
|
Transverse (adj.) (Reitsma, 1970) | ||
Synonym of
oblate.
|
Transversal furrow (Iversen and Troels-Smith, 1950) | ||
Synonym of
endocolpus. |
Trema (pl. tremata) (Erdtman and Straka, 1961) | ||
Synonym of
aperture.
|
-treme (Erdtman and Straka, 1961) | ||
A suffix synonymous with
aperture.
|
Tri- | ||
A prefix for three. |
Trichotomocolpate (adj.) (Erdtman, 1945a) | ||
Describing a
pollen grain with a three-armed
colpus. Example: Trapa natans (Trapaceae). |
Trichotomosulcate (adj.) (Erdtman, 1952) | ||
Describing a
pollen grain with a three-armed
sulcus. Example: Elaeis guineensis (Palmae). |
Tricolpate, tricolporate, triporate (adj.) (Iversen and Troels-Smith, 1950) | ||
Describing
pollen grains with three
ectocolpi, three
compound apertures or three
pores.
|
Trifolium (pl. trifolia) (Potonié, 1956) | ||
A three-bladed,
proximal feature arising from the
proximal pole of a
megaspore. Example: Capulitriletes.
|
Trilete (adj.) (Erdtman, 1943) | ||
Describing a spore with three laesurae, thus showing a trilete mark. Example: Pteridium
(Hypolepidaceae).
|
Trilete mark | ||
The triradiate mark of a
trilete spore. |
Triprojectate (adj.) (Mtchedlishvilli, 1961) | ||
Describing
projectate pollen
with three projecting arms on which the apertures
are situated, as for example in the Triprojectacites group of fossil pollen grains. Example:
Aquilapollenites. |
Tryphine (Erdtman, 1969) | ||
A material deposited on the surface of pollen grains by the breakdown of the tapetum and differing from
pollenkitt in that it contains membraneous components derived from organelles.
|
Tuberculate (adj.) (Moar, 1993) | ||
Covered with knobbly projections. |
Tubulus (pl. tubuli) (Erdtman, 1952) | ||
A general term for a small channel, applied in palynology to a channel through the nexine. |
Tula (pl. tulae, adj. tulate) (Jansonius and Pocock, 1969) | ||
A sexinal inflation in gymnospermous pollen at the end of the axis of the
distal sulcus or
leptoma. Example:
Ovalipollis. |
Tumescence (pl. tumescences) (Batten and Christopher, 1981) | ||
A gradual increase in thicknes of the wall layer(s) from
a point in the equatorial interradial region to the
germinal aperture. Example:
Megatriopollis.
|
Turma (pl. turmae) (Potonié, 1956) | ||
An artificial suprageneric grouping of form-genera of fossil
spores and
pollen.
|
Turriform (Balme, 1988) | ||
Biform sculptural elements consisting of a capitate basal portion surmounted by a sharply contracted distal spine. Example: Dibolisporites. |
Ubisch body (pl. ubisch bodies) (Rowley, 1963) | ||
A distinctive, orbicular granule of
sporopolleninsporopollenin produced by the
tapetum, particularly in plants with secretory tapeta.
|
Ulculus (pl. ulculi, adj. ulculate) (Walker and Doyle, 1975) | ||
A rounded
ectoaperture not situated at a
pole. Example: Poaceae.
|
Ulcus (pl. ulci, adj. ulcerate) (Erdtman, 1952) | ||
A rounded
ectoaperture situated at the
distal or proximal
pole of a pollen grain. Examples:
Sparganium (Sparganiaceae), Typha (Typhaceae).
|
Uniplanar tetrad (Walker and Doyle, 1975) | ||
A tetrad in which the individual members lie more or less in one
plane.
|
Urceolate (adj.) (Ferguson et al., 1983) | ||
Describing a type of ornamentation consisting of urn-
shaped elements situated on the footlayer. Example: Pinanga aristata (Palmae). |
Vacuoles (Grebe, 1971) | ||
Rounded to elongated spaces within an equatorial faeture.
Example: Vallatisporites ciliares.
|
Vallum (pl. valla) (Iversen and Troels-Smith, 1950). | ||
Synonym of
murus.
|
Valva (pl. valvae, adj. valvate) (Potonié and Kremp, 1955) | ||
Radial thickenings in the areas beyond the ends of the
laesurae of
trilete spores. Example:
Triquitrites tribullatus.
|
Velum (pl. vela, adj. velate) (Thomson and Pflug, 1953) | ||
A feature of a
monosaccate
pollen grain in which the
saccus is convoluted. Example: Tsuga (Pinaceae). |
Vermiculate (adj.) (Kosanke, 1950; Harris, 1955) | ||
A general descriptive term used to describe winding
features.
|
Verruca (pl. verrucae, adj. verrucate) (Iversen and Troels-Smith, 1950) | ||
A wart-like sexine element, more than 1µm wide,
that is broader than it is high and is not constricted at the base. Example: Plantago
(Plantaginaceae). |
Verrucose (adj.) (Erdtman, 1969) | ||
Synonym of verrucate. |
Vesiculate (adj.) (Iversen and Troels-Smith, 1950) | ||
Synonym of saccate. |
Vestibulum (pl. vestibula) (Potonié, 1934) | ||
A separation between layers of the exine forming a cavity between the inner and outer pores. For example: Betula (Betulaceae).
|
Viscin thread (pl. viscin threads) | ||
A general botanical term (Jackson, 1928), applied in
palynology for an
acetolysis resistant,
sporopollenin thread arising from the
exine of a pollen grain,
usually from the proximal surface. Example:
Oenothera (Onagraceae). |
Y-mark (Potonié, 1934) | ||
Synonym of
trilete mark. |
Z-layer (Heslop-Harrison, 1979) | ||
Synonym of exintine. |
Zona (pl. zonae, adj. zonate) (Potonié and Kremp, 1955) | ||
A thin outer structure of a
spore that projects at the
equator, but does not extend over the
distal face or proximal
face. Example: Cirratriradites saturni.
|
Zona- (adj. zonate) (Walker and Doyle, 1975) | ||
Prefix indicating ring-like.
|
Zona-aperturate (adj.) (Walker and Doyle, 1975) | ||
Describing a
pollen grain with a ring-like aperture.
|
Zonasulculus (Walker and Doyle, 1975) | ||
A ring-like sulculus around the equator. Example: Nymphaea violacea
(Nymphaeaceae).
|
Zonasulcus (Walker and Doyle, 1975) | ||
A meridional ring-like
sulcus perpendicular to the equator. Example: Laurelia
novaezelandiae (Monimiaceae).
|
Zoni- (sensu Erdtman and Vishnu-Mittre, 1956) | ||
Synonym of zono-. |
Zoni- (sensu Walker and Doyle, 1975) | ||
Prefix indicating a latitudinal orientation. |
Zono- (Erdtman and Straka, 1961) | ||
A prefix indicating features located equatorially.
|
Zonoaperturate (adj.) (Erdtman and Straka, 1961) | ||
Describing a
pollen grain with
apertures situated only at the
equator.
|
Zonorate (adj.) (Erdtman, 1952) | ||
Describing a
pollen grain with a continuous endoaperture (os) around the equator.
|
Zwischenkörper (pl.) (Fritzsche, 1837) | ||
A lens-shaped body below the
aperture of certain unacetolysed
pollen grains.
|